In this photo of early Ushak Medallion border from his book "L'Ushak" Alberto Boralevi ( notes the Kufic interlace combined with floral motifs.  Like the medallion arabesques described previously, this seems to connect the curvilinear large format Ushak designs with their predecessors, the Lotto and Holbein, as well as with Timurud and Seljuk carpets.

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